Whole plant biology of fossil plants and their use in phylogenetic analysis.
Evolution of ferns, gymnosperms, and aquatic angiosperms.
Phylogenetic trends in the Coniferales.
Fossil record of monocots.
Reproductive biology of the Araucariaceae and Podocarpaceae, including fossil conifers.
Anatomy and morphology of vascular plants.
Cretaceous and Tertiary plants: ferns, gymnosperms, angiosperms.
Plant/fungal interactions in the fossil record/fossil mycorrhizae.