Exploration of key concepts regarding the ecology of forest ecosystems at varying temporal and spatial scales. Emphasis will be on relationships between biotic and abiotic components of the ecosystem. Topics covered will include flows of energy and matter, ecosystem dynamics, forest landscapes and biodiversity, impacts of natural and anthropogenic disturbance, forest conservation and ecosystem management. Lab exercises during the first month are held outside. May require payment of additional student instructional support fees. Refer to the Tuition and Fees page in the University Regulations section of the Calendar. Not to be taken if credit received for REN R 322.