The course will introduce students to theory and techniques employed in the analysis of physical, hydrological, chemical, and ecological properties of ecosystems using a watershed (catchment) approach. Focus will be on landscape approaches relating interactions or linkages between upland, wetland/riparian, and surface-water in the study of the natural ecohydrologic function and response to disturbance of watershed ecosystems. Emphasis will be placed on Boreal Alberta. Topics are covered through reading the literature and group discussions. Seminars are the same as for BIOL 440, but with additional assignments and evaluation appropriate to graduate studies. Prerequisite: consent of instructor. Credit cannot be obtained for both BIOL 440 and 540.